How to change the order in which users are displayed in admin?

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Nope, the line you found seems to be some basic preliminary query. Selection of users for display in table seems to be handled by WP_User_Search class. Luckily it can be hooked into easily.

Try this (changes order to descending):

add_action('pre_user_search', 'change_user_order');

function change_user_order($query) {

    $query->query_orderby = ' ORDER BY user_login DESC';
}

PS I am not sure if this can interfere with something else, so might require checks to only run on that specific page.

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