I solved this with help from the main StackOverflow site. One of the commenters suggested that the open_basedir directive might be set in php.ini – and that was the issue. Here is the open_basedir setting:
open_basedir = "/var/www/vhosts/lifering.org/:/tmp/"
I created a hard link mysql (could have been a symlink) to /usr/bin/mysql in the /tmp directory and then changed the MySQL path statement in WP-DBManager to read /tmp/mysql. The error message cleared immediately.
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