Answered by @Vino here: https://stackoverflow.com/questions/12066316/sub-folder-css-template-headers-that-are-read-by-admin-panel/12139043#comment16259096_12139043
“File wp-includes/class-wp-theme.php, line 191, WP 3.4.1 source. but… there are no filters, and the class is final. You can edit at your peril, ’cause updates to WP will overwrite your efforts.”
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