The difference between ?= and ?! is that the former requires the given expression to match and the latter requires it to not match. For example a(?=b) will match the “a” in “ab”, but not the “a” in “ac”. Whereas a(?!b) will match the “a” in “ac”, but not the “a” in “ab”.
The difference between ?: and ?= is that ?= excludes the expression from the entire match while ?: just doesn’t create a capturing group. So for example a(?:b) will match the “ab” in “abc”, while a(?=b) will only match the “a” in “abc”. a(b) would match the “ab” in “abc” and create a capture containing the “b”.
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