Getting Access Denied when calling the PutObject operation with bucket-level permission

To answer my own question:

The example policy granted PutObject access, but I also had to grant PutObjectAcl access.

I had to change

"s3:PutObject",
"s3:GetObject",
"s3:DeleteObject"

from the example to:

"s3:PutObject",
"s3:PutObjectAcl",
"s3:GetObject",
"s3:GetObjectAcl",
"s3:DeleteObject"

You also need to make sure your bucket is configured for clients to set a public-accessible ACL by unticking these two boxes:

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