I think the problem is here
>] <h3 id="comments-title"><?php
printf( _n( 'One Response to %2$s', '%1$s Responses to %2$s', get_comments_number() ),
number_format_i18n( get_comments_number() ), '<em>' . get_the_title() . '</em>' );
?></h3>
Try changing that to
<h3 id="comments-title"><?php
printf( _n( 'One Response to %2$s', '%1$s Responses to %2$s', number_format_i18n( get_comments_number() ), '<em>' . get_the_title() . '</em>' );
?></h3>
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