This line has wrong syntax:
<?php if ( is_singular() ) wp_enqueue_script ( 'comment-reply'); wp_head() ; ?>
The call to wp_head() should not be on the same line as the conditional for the comment-reply script.
Change it to this:
<?php
// Conditional to determine if comment-reply form script should be included
if ( is_singular() ) wp_enqueue_script ( 'comment-reply');
// Fire the `wp_head` hook, which should happen always, on every pageload
wp_head();
?>
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